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(B) MV and MP females also show a greater preference for social investigation as they spend more time investigating a novel conspecific than an inanimate object (mean±SEM) (n = 10 per group). (C and D) MO, MV, and MP males, which do not display enhanced oligodendrogliogenesis, show no difference in performance on the horizontal ladder rung. MO, MV, and MP males all demonstrate a greater preference for investigating the novel conspecific versus the inanimate object (mean±SEM) (n = 8 per group). (E) MO females sired by MV or MP mothers demonstrate less slipping on the horizontal ladder rung over time (mean±SEM) (n = 10 per group). (F) MO females sired by MV or MP mothers display greater investigation towards a novel conspecific (mean±SEM) (MO females (P: MO female×MO male) (n = 10), MO females (MV female×MO male) (n = 11) and MO females (MP female×MO male) (n = 13)). info:doi/info:doi/10.1371/journal.pone.0062701.g004 Public Library of Science Figure 4 Creative Commons Attribution License info:doi/10.1371/journal.pone.0062701 C       C " l  !1 "Av789Qu#2VWaqw356BSTXrst$Rb4U%CE'FcK  !1AQa2q"r3BRSb#4TUCcs ?2东76fZb:s)~}Ө$ }W<&f쁕{c3FBdk"M&sqɨ rbCq/SXݹZƜ0㧑)Iʽ58iTp(ȳI:k<5DUK5N.l 5>ZD1k($02UWz750hQCnRRRRRRRRRRRRRRRRRRRUOfUWVq/n0#?L71dɡC{J+:xD=W%nEes>LQ]6Z1Eg]dꪢfPR]kB@ =CA2eMԻw,hZq_ \q4SQr(WiG@u4GM9lb}jET`j┥E 1u=n8,=i'>bEiu àwkA#fyX/ή8LXȡ 0+8EncP$J3R\ƷYb$eLPQ]P! : ""BTg-Ij/A6O훺MzRqDI":@u)0y:si@)@PIyC0)k ǙhO^zWa_YIq$MtAbU31G@Dyq>3ouf^RQ'HGM]&FDB bkV-Z^<79fmع8vC*&3XXt BD _vͭpp%8k$˶TRHuE=!P %5XĶ- yn Er(seP۔UE1=9G@((\C\VPq8Nkd rifcH̊$MCMB cܣo0ƭ@)@Qɽ@!UQPB S {@tkXῈ>'sZv7i%ǘ˱;J ʸxA@US'СJP)JP)JP)JP)J.+ ҊVrILR(@ = ڔ[E)-2eNLX3<"#eҫ>1O>1WGo(:#+Xtս-k݊VmTPQDuЪh] HYTHJs[ OnS 6]ܲ:>G&+q<'a͏QHˆj)'0T㏳V˱=oZg8*ԅgӔirn00_1GH7 L'0iGO> O4AC[xFE96U""]zZ6z13w~8,q೵_T.t\&dWHj@:g#/0՗}c=y>x~ڳ#_BG-#]*&A2 (:p7T$@9RxW\_*sg^!gB4aR!JpCpq\'bۊ]줓r냮SCDh.84$ jBչ-njvr*'+T u v뮚s,7#}:x}Y2 K(8 r cѲ_@&夙@.LDt(j"#R\\=1ּKu;a8\"( n P{-km,˓hBN Us9AOT: QDpj |͙`=%Gw,l#l.`C+)4nR 䊩(ja&qk[ݫ ܋Y6r3L(q a DɏRz{scKgȵko}ahe2ّ<K][JƄ h\8]nLJr*Gr nlLb6EwLdٵ˿nU fzJr&܍H&'01s8kk.ː{8yCVQETT4Є|q63)SLʲYd>x .nI2!Y@CkmڶVﳧZ9-&yȴHMQp":j *b%|=!){(DV ._0OnP]e (l2۠ C4peSWxBZڶ~Gr-?`\[n6ZM2!AN@ A4Gǐw.R,ƠYb%rX5;b (WH"Hm@)@)@)@)A޶m6㜪r/֨5 9D A ?7uvα!$,/!왪LmP r'9`\АBFBVPED!1 SDMD;#{eE\RL=Qgp=u{@ Qrc~qЏaż\SٴLn2I4)RJhȥ((*0}9iBV}~Qڣ|ƛ%k'qGiZe¯s9|ɜ{eޘg> ą17 !M cD2'_>v|c`c3diI0_~S|ɢ ӺʑDǖ@k?R?Ov}O%]X-*7Qi[3U:AD7b9aZGWAޮePT(v$D*nn!nBaDa!e3z\ $\P=SXAEDY2!L"#}{`&Zr㆔zБ`'ADrPSnO `Һ^1(G2,M`EHєsa0) dp9a1L%4Jsw$ɿвד!)47UM]5P)9Gl7։CQ5)iI},qi[ĹLM  YRL Tj>JJ)nE͢"gLשD6}o^/n,T `kD8UK VLѪqQA~LD hmż.krƈg|7s岓nDfi+c'ХnA8z*OJM i(r6t %1!S1PSxs`2+K}%2ಪ:U vȦ|I0ˆ~%ǩ̢TE'^$zET90'QM ͨP/5#Wq"9&VL\!"nIC*E}LcU{ݗ36FNلo+(|E]6D)s)V"PHqÖ&q  3HڀKs6JAnm(C ǧ(cܿjDҭܳB!I@LBJR]EjH5c3.]3;Q_h9L:SpWoUWZw:v174#wmZLc+$@DK%-7-|܉(D@[/_ [M7YeFI-~pJ k"!.B1uF0xs`[:O@Q6kB:5 BMt7mhTj2%Y@߱Yɜ<AҔRRTk!d{'[j^}hdVI,C4ΡvˆSPD@ZG|ZL>b˒bmq[:M36!{P^$Tl$BUr8MQ\XP )DD`P6(V6mGCHɎщ̙ r6LJ!APJ8t;6T;e*/ݘ;VE0HfD@AS( CAh5Rkb12˘/G\S!`}GJ"a"ƪ_ӕ!Lp(0.odgŶ+n8 SydQ䮄L .$>$צԠҰ3V>b8LugM,HET@>6JVQNsV RU|㗗nb%Orpj1pMM]JjAv¡m!^n5AA: sG:>mе&,xɎv&:fKƹS¶b.5Ch-}z~A: !CŨ>Jtr1 M-@@ө :㼣 IlXˡHch >/NA:iz BWY˧m_Jؚ[.wODtP.'SCJ۽'lgeL\Qv 'P"͕IQH9p b8Dk$㏠GŎ-N &Q:-g-Mb*AQT1JUmILQ*R R>4ߡ+]>ӌM?(qQ>cM8nA~ҔRRRRRRpdg 5dY>1톂])f) 5@-?\WDŽK?L"TF<%hHVAdYM9Ch:t Fy?3|y)yGIP"nbLuMzک Ѡ9r/8Uw 0UUU8&1=DDDDDjOQojUUcbs|䋳\Dzem릺ע R R dLobCsg͵;ꭔ(z m+z^ح1ZޕYxGcyfl,[ɇ)_.zt[h z ӡo Csk-C5U-bq 5KK4HG[m5o)EId|;޹_˜( @@zW)J)J)J)J)J)J)J)J)J)J)J)JKir㛪2H- sJ5P9 mDC+Hٱ׍LF꾶%- Cw)K` z4QdpٙVR-)imw,T&,B3Qda ’`eG)G@ ֕ߌ2 wruiȴUKE37PPU1CT .tdEb9D5Aҫac,CnY U3VŔT1RD*DS/@މg/[.ȶc=t\SI$ M>16:P\/W>og3ˁ)>M,R~ap-"Nq 5#6ߐC!a7cev*̓XB P c 41 Zx bo7QFk<`{r6)uizOY.[4;8}A!\ :.hkCAi&uj ^m.E3QxL}%DhSOq VTSghz}7:TOYoz2u,RR?t F eѯ߄HʅZ5Wf/מ"D~T(=)@)@)@)@)A8,c"Ymks&8fr&@QT :nFOovStnM i7kb4A.Dʼi'S&\eꘉ8*b/, C1l.\e&ۗqMܶV:m!M5c!>ޚiA}W]qm%k\,\EXn'1d B `(t҂86Ӆ q$bw"I:Iv9T)vNR ] PcK[D1$(QH9YCnQU8cDtǹG;a[ QNsV RWZn4;y_!fX ]wvӦUImGC]1ғE%Ath#Ap];])Ju7EyC/p+p1 !LA YቈT=qT-;VvfJwNSw;A7MV"ȪP:jb;¸0W [Y)Ghyq_r\hQvITͩL\ڳY6^FȮnEZ3M[ΕY(Je2`>ֱmɹĹ:<|k431e,W7.Sd=42Ч2z*il WͻtI `M2n8vh־ R R R R_y'ZoᓗR5`yy$ڨ 5*52ZOˣŶ]Q ]JP)JP)JP)JP)JPWceʠuFZR7 <b)&n͖q8ָxz!X<\?Ȩl̂)ULT ˮL-K匼vhp[~S3rmMM+ZrRu ŠI.FǷ\LͰ!P h_ -$a3+{Jxi%[k&9*8ME!ʖ%L1DL":h:Pf;τw("TZnlm Q&f)UΩA6+EP?x^< 4|,i$n#,{4HK!D(W+i&uj1 Zx bo7QFiJP)JP)JP)JP)JP)JP)JPy1S?wK?L"TFU?wZ5lUO?]&~ت~;ѓeD/Uw'R)J)J)H.o0|k_.( &_-*_<=DUUvby$GBJR R R R R+3 HK+w]t鑔/5Url}ޘ#HQrd{q$[XY9Pt"xdE+sQ@9:(t/ܮ80<:-gpum W\cRQh4=q%c`NLA|sureCzJP Tp]"(( LQ (3ؚcY2}oZ-"mG2nNd;d9|C{yAD\Uwa\,tg6Ts*^BHmzzJ]QZ5wt5oH+U B8{/Ԏr ʡ@`ӥ eub6\}-g{Z鋷 L`UA0^XB(m(x[9aWvo~SoٓDy[:iO Hp"Іz)@)@m̾t 2ׅ+'ciԟM5u:t]V0S(gEkUoP! 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