Myotinae is one of the four subfamilies of Vespertilionidae, itself one of twenty families of bats in the mammalian order Chiroptera and part of the microbat suborder. A member of this subfamily is called a myotine, or a mouse-eared bat. They are found in all continents besides Antarctica, primarily in forests and caves, though some species can also be found in shrublands, grasslands, rocky areas, or deserts. They range in size from the Taiwan broad-muzzled bat, at 3 cm (1 in) plus a 3 cm (1 in) tail, to the large myotis, at 10 cm (4 in) plus a 6 cm (2 in) tail. Like all bats, myotines are capable of true and sustained flight, and have wing lengths ranging from 2 cm (1 in) to 7 cm (3 in). They are all insectivorous and eat a variety of insects and spiders,[1] but some bats with long toes which trawl for insects on top of the water, such as the long-fingered bat Daubenton's bat, Maluku myotis, Rickett's big-footed bat, and pond bat, may sometimes supplement their diet with small fish from still waters. The fish-eating bat also regularly eats fish and crustaceans as well as insects, and is the only bat species that hunts fish in the ocean.[2] Almost no myotines have population estimates, though seven species—the Atacama myotis, eastern small-footed myotis, Findley's myotis, flat-headed myotis, frosted myotis, little brown bat, and peninsular myotis—are categorized as endangered species, and two species—the Nimba myotis and Yanbaru whiskered bat—are categorized as critically endangered.

A photograph of a greater mouse-eared bat hanging upside-down. It is gray and small.
Greater mouse-eared bat (Myotis myotis)

The 121 extant species of Myotinae are divided between three genera: Eudiscopus and Submyotodon with one species each, and Myotis, or the mouse-eared bats, with the other 119. A few extinct prehistoric myotine species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.[3]

Conventions

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IUCN Red List categories
Conservation status
 EX Extinct (0 species)
 EW Extinct in the wild (0 species)
 CR Critically Endangered (2 species)
 EN Endangered (7 species)
 VU Vulnerable (8 species)
 NT Near threatened (11 species)
 LC Least concern (65 species)
Other categories
 DD Data deficient (28 species)
 NE Not evaluated (0 species)

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the myotine's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted. Population figures are rounded to the nearest hundred.

Classification

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Myotinae, one of the four subfamilies of the family Vespertilionidae, contains 121 extant species divided into 3 genera.

Subfamily Myotinae

  • Genus Eudiscopus (disk-footed bat): one species
  • Genus Myotis (mouse-eared bats): 119 species
  • Genus Submyotodon (Taiwan broad-muzzled bat): one species

Myotines

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The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.[4]

Genus Eudiscopus Conisbee, 1953 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Disk-footed bat


E. denticulus
(Osgood, 1932)
Southeastern Asia
 
Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Forest[6]
 LC 


Unknown  [6]

Genus Myotis Kaup, 1829 – 119 species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Alcathoe bat

 

M. alcathoe
von Helversen & Heller, 2001
Europe and Turkey
 
Size: 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[7]

Habitat: Forest[8]
 DD 


Unknown  [8]

Anjouan myotis


M. anjouanensis
Dorst, 1960
Madagascar
 
Size: About 5 cm (2 in), plus about 5 cm (2 in) tail
about 4 cm (2 in) arm/wing length[7]

Habitat: Unknown[9]
 DD 


Unknown  [9]

Anna Tess's bat


M. annatessae
Kruskop & Borisenko, 2013
Vietnam and Laos
 
Size: 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[10]

Habitat: Forest[11]
 DD 


Unknown  [11]

Annamit myotis


M. annamiticus
Kruskop & Tsytsulina, 2001
Vietnam
 
Size: About 4 cm (2 in), plus about 3 cm (1 in) tail
about 4 cm (2 in) arm/wing length[10]

Habitat: Forest and inland wetlands[12]
 DD 


Unknown  [12]

Arizona myotis


M. occultus
Hollister, 1909
Mexico and southwestern United States
 
Size: 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[5]

Habitat: Forest, rocky areas, caves, and desert[13]
 LC 


Unknown  [13]

Atacama myotis


M. atacamensis
Lataste, 1892
Chile and Peru
 
Size: 4–5 cm (2–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Shrubland, rocky areas, and desert[15]
 EN 


Unknown  [15]

Australian myotis


M. australis
Dobson, 1878
Australia
 
Size: Unknown[16]

Habitat: Unknown[16]
 DD 


Unknown  [16]

Barbados myotis


M. nyctor
LaVal & Schwartz, 1974
Lesser Antilles in the Caribbean Size: 4–5 cm (2–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest and caves[17]
 VU 


Unknown  [17]

Bechstein's bat

 

M. bechsteinii
(Kuhl, 1817)
Europe and western Asia
 
Size: 3–6 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest, shrubland, and caves[19]
 NT 


Unknown  [19]

Beijing mouse-eared bat


M. pequinius
Thomas, 1908
Eastern China
 
Size: About 6 cm (2 in), plus about 4 cm (2 in) tail
4–6 cm (2–2 in) arm/wing length[18]

Habitat: Forest and caves[20]
 LC 


Unknown  [20]

Black myotis

 

M. nigricans
Schinz, 1821

Four subspecies
  • M. n. carteri
  • M. n. extremus
  • M. n. nigricans
  • M. n. osculatii
Mexico, Central America, and South America
 
Size: 3–6 cm (1–2 in), plus 2–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[7]

Habitat: Forest, savanna, shrubland, and caves[21]
 LC 


Unknown  [21]

Bocharic myotis


M. bucharensis
Kuzyakin, 1950
Central Asia
 
Size: 4–6 cm (2–2 in), plus 4–6 cm (2–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Caves[22]
 DD 


Unknown  [22]

Bornean whiskered myotis


M. borneoensis
Hill & Francis, 1984
Island of Borneo (in green)
 
Size: Unknown length, plus 4–5 cm (2–2 in) tail
4–5 cm (2–2 in) arm/wing length[10]

Habitat: Forest[23]
 DD 


Unknown  [23]

Brandt's bat

 

M. brandtii
(Eversmann, 1845)

Two subspecies
  • M. b. brandtii
  • M. b. gracilis
Europe and western Asia (in red)
 
Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[7]

Habitat: Forest, shrubland, grassland, inland wetlands, and caves[24]
 LC 


Unknown  [24]

Burmese whiskered myotis

 

M. montivagus
(Dobson, 1874)
Southern Asia (in leftmost red)
 
Size: Unknown length
3–5 cm (1–2 in) arm/wing length[10]

Habitat: Forest and caves[25]
 DD 


Unknown  [25]

California myotis

 

M. californicus
(Audubon & Bachman, 1842)

Four subspecies
  • M. c. californicus
  • M. c. caurinus
  • M. c. mexicanus
  • M. c. stephensi
Western North America
 
Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Forest, caves, and desert[26]
 LC 


Unknown  [26]

Cape hairy bat


M. tricolor
(Temminck, 1832)
Sub-Saharan Africa
 
Size: 5–7 cm (2–3 in), plus 3–6 cm (1–2 in) tail
4–6 cm (2–2 in) arm/wing length[7]

Habitat: Forest, savanna, and shrubland[27]
 LC 


Unknown  [27]

Cave myotis

 

M. velifer
(Allen, 1890)

Five subspecies
  • M. v. brevis
  • M. v. grandis
  • M. v. incautus
  • M. v. magnamolaris
  • M. v. velifer
United States and Mexico
 
Size: 4–6 cm (2–2 in), plus 3–6 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[14]

Habitat: Forest and caves[28]
 LC 


Unknown  [28]

Chilean myotis

 

M. chiloensis
(Waterhouse, 1840)
Southern South America
 
Size: 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[14]

Habitat: Rocky areas and forest[29]
 LC 


Unknown  [29]

Chinese water myotis


M. laniger
Peters, 1870
Eastern Asia
 
Size: 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[10]

Habitat: Forest and caves[30]
 LC 


Unknown  [30]

Cinnamon myotis


M. fortidens
Miller & Allen, 1928

Two subspecies
  • M. f. fortidens
  • M. f. sonoriensis
Guatemala and Mexico
 
Size: 4–6 cm (2–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Forest and caves[31]
 LC 


Unknown  [31]

Cryptic myotis

 

M. crypticus
Ruedi, Ibáñez, Salicini, Juste, & Puechmaille, 2019
Southern Europe Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest, inland wetlands, and caves[32]
 NT 


Unknown  [32]

Csorba's mouse-eared bat


M. csorbai
Topál, 1997
Nepal Size: Unknown[33]

Habitat: Forest[33]
 DD 


Unknown  [33]

Curacao myotis


M. nesopolus
Miller, 1900

Two subspecies
  • M. n. larensis
  • M. n. nesopolus
Northern South America
 
Size: About 3 cm (1 in), plus about 4 cm (2 in) tail
2–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest and shrubland[34]
 LC 


Unknown  [34]

Dark-nosed small-footed myotis


M. melanorhinus
Merriam, 1890
Western North America
 
Size: 3–5 cm (1–2 in), plus 2–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Caves and forest[35]
 LC 


Unknown  [35]

Daubenton's bat

 

M. daubentonii
(Kuhl, 1817)

Six subspecies
  • M. d. chasanensis
  • M. d. daubentonii
  • M. d. loukashkini
  • M. d. nathalinae
  • M. d. ussuriensis
  • M. d. volgensis
Europe and Asia
 
Size: 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest, shrubland, inland wetlands, rocky areas, caves, and neritic marine[36]
 LC 


Unknown  [36]

David's myotis


M. davidii
Peters, 1869
Eastern Asia
 
Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[7]

Habitat: Caves and forest[37]
 LC 


Unknown  [37]

Diminutive bat


M. diminutus
Moratelli & Wilson, 2011
Ecuador and Colombia Size: About 4 cm (2 in), plus about 3 cm (1 in) tail
about 3 cm (1 in) arm/wing length[14]

Habitat: Forest[38]
 DD 


Unknown  [38]

Dinelli's myotis

 

M. dinellii
Thomas, 1902
Southern South America (in green)
 
Size: 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest[39]
 LC 


Unknown  [39]

Dominican myotis


M. dominicensis
Miller, 1902
Dominica and Guadeloupe in the Caribbean
 
Size: 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Inland wetlands and caves[40]
 VU 


Unknown  [40]

Eastern long-fingered bat

 

M. macrodactylus
(Temminck, 1840)

Three subspecies
  • M. m. continentalis
  • M. m. insularis
  • M. m. macrodactylus
Eastern Asia
 
Size: 4–8 cm (2–3 in), plus 2–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest, inland wetlands, and caves[41]
 LC 


Unknown  [41]

Eastern small-footed myotis

 

M. leibii
Audubon & Bachman, 1842
Eastern United States and Canada
 
Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Caves, rocky areas, and forest[42]
 EN 


Unknown  [42]

Eastern water bat

 

M. petax
Hollister, 1912
Eastern Asia
 
Size: 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[18]

Habitat: Caves, inland wetlands, and forest[43]
 LC 


Unknown  [43]

Elegant myotis


M. elegans
Hall, 1962
Mexico and Central America
 
Size: 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest[44]
 LC 


Unknown  [44]

Escalera's bat

 

M. escalerai
A. Cabrera, 1904
Southwestern Europe Size: 4–5 cm (2–2 in), plus 4–5 cm (2–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest, shrubland, and caves[45]
 NT 


Unknown  [45]

Far Eastern myotis


M. bombinus
Thomas, 1906

Two subspecies
  • M. b. amurensis
  • M. b. bombinus
Eastern Asia
 
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest and caves[46]
 NT 


Unknown  [46]

Felten's myotis

 

M. punicus
Felten, 1977
Northern Africa and Mediterranean islands
 
Size: 6–8 cm (2–3 in), plus tail
5–7 cm (2–3 in) arm/wing length[18]

Habitat: Caves, grassland, shrubland, and forest[47]
 DD 


Unknown  [47]

Fish-eating bat

 

M. vivesi
Ménégaux, 1901
Western Mexico
 
Size: 8–9 cm (3–4 in), plus 5–8 cm (2–3 in) tail
5–7 cm (2–3 in) arm/wing length[14]

Habitat: Rocky areas and caves[48]
 VU 


Unknown  [48]

Findley's myotis


M. findleyi
Bogan, 1978
Tres Marías Islands west of Mexico
 
Size: 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail
2–4 cm (1–2 in) arm/wing length[5]

Habitat: Unknown[49]
 EN 


Unknown  [49]

Flat-headed myotis

 

M. planiceps
Baker, 1955
Central Mexico
 
Size: 3–6 cm (1–2 in), plus 2–3 cm (1–1 in) tail
2–3 cm (1–1 in) arm/wing length[5]

Habitat: Forest and caves[50]
 EN 


240  [50]

Fraternal myotis

 

M. frater
(Allen, 1923)
Eastern Asia (in red)
 
Size: 3–5 cm (1–2 in), plus 4–6 cm (2–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest and caves[51]
 LC 


Unknown  [51]

Fringed long-footed myotis


M. fimbriatus
Peters, 1870
Eastern Asia
 
Size: 3–6 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Caves[52]
 LC 


Unknown  [52]

Fringed myotis

 

M. thysanodes
Miller, 1897

Four subspecies
  • M. t. aztecus
  • M. t. pahasapensis
  • M. t. thysanodes
  • M. t. vespertinus
Western North America
 
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
4–5 cm (2–2 in) arm/wing length[5]

Habitat: Forest, rocky areas, caves, and desert[53]
 LC 


Unknown  [53]

Frosted myotis


M. pruinosus
Yoshiyuki, 1971
Japan
 
Size: 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[10]

Habitat: Forest and caves[54]
 EN 


Unknown  [54]

Geoffroy's bat

 

M. emarginatus
(Geoffroy, 1806)

Three subspecies
  • M. e. desertorum
  • M. e. emarginatus
  • M. e. turcomanicus
Europe, northern Africa, and western Asia
 
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[7]

Habitat: Caves, grassland, and shrubland[55]
 LC 


Unknown  [55]

Gomantong myotis


M. gomantongensis
Francis & Hill, 1998
Island of Borneo in Malaysia
 
Size: Unknown length, plus 3–5 cm (1–2 in) tail
4–5 cm (2–2 in) arm/wing length[10]

Habitat: Caves[56]
 LC 


Unknown  [56]

Gray bat

 

M. grisescens
Howell, 1909
Eastern United States
 
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
4–5 cm (2–2 in) arm/wing length[14]

Habitat: Forest and caves[57]
 VU 


Unknown  [57]

Greater mouse-eared bat

 

M. myotis
(Borkhausen, 1797)

Two subspecies
  • M. m. macrocephalicus
  • M. m. myotis
Europe and western Asia
 
Size: 6–9 cm (2–4 in), plus 4–6 cm (2–2 in) tail
5–7 cm (2–3 in) arm/wing length[18]

Habitat: Forest, shrubland, and caves[58]
 LC 


Unknown  [58]

Guatemalan myotis


M. cobanensis
Goodwin, 1955
Guatemala
 
Size: About 4 cm (2 in), plus about 4 cm (2 in) tail
about 4 cm (2 in) arm/wing length[14]

Habitat: Unknown[59]
 DD 


Unknown  [59]

Hairy-faced bat


M. annectans
Dobson, 1871
Southern and southeastern Asia
 
Size: 5–7 cm (2–3 in), plus 4–5 cm (2–2 in) tail
4–5 cm (2–2 in) arm/wing length[10]

Habitat: Forest[60]
 LC 


Unknown  [60]

Hairy-legged myotis

 

M. keaysi
Allen, 1914

Two subspecies
  • M. k. keaysi
  • M. k. pilosotibialis
Mexico, Central America, and northern and western South America
 
Size: 5–7 cm (2–3 in), plus 2–5 cm (1–2 in) tail
Unknown arm/wing length[5]

Habitat: Forest[61]
 LC 


Unknown  [61]

Herman's myotis


M. hermani
Thomas, 1923
Indonesia
 
Size: 5–6 cm (2–2 in), plus 3–6 cm (1–2 in) tail
4–6 cm (2–2 in) arm/wing length[7]

Habitat: Forest[62]
 DD 


Unknown  [62]

Himalayan whiskered bat


M. siligorensis
Horsfield, 1855

Four subspecies
  • M. s. alticraniatus
  • M. s. siligorensis
  • M. s. sowerbyi
  • M. s. thaianus
Central, eastern, and southeastern Asia
 
Size: Unknown length
about 3 cm (1 in) arm/wing length[10]

Habitat: Forest and caves[63]
 LC 


Unknown  [63]

Hodgson's bat

 

M. formosus
(Hodgson, 1835)
Central and eastern, and southeastern Asia Size: 5–6 cm (2–2 in), plus 3–6 cm (1–2 in) tail
4–6 cm (2–2 in) arm/wing length[7]

Habitat: Forest, shrubland, grassland, and caves[64]
 NT 


Unknown  [64]

Horsfield's bat

 

M. horsfieldii
Temminck, 1840

Five subspecies
  • M. h. deignani
  • M. h. dryas
  • M. h. horsfieldii
  • M. h. jeannei
  • M. h. peshwa
Southern and southeastern Asia
 
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[10]

Habitat: Forest and caves[65]
 LC 


Unknown  [65]

Ikonnikov's bat


M. ikonnikovi
Ogniov, 1912
Eastern Asia
 
Size: 3–6 cm (1–2 in), plus 2–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[7]

Habitat: Forest, inland wetlands, and caves[66]
 LC 


Unknown  [66]

Indiana bat

 

M. sodalis
Miller, 1898
Eastern United States
 
Size: 4–5 cm (2–2 in), plus 2–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Forest and caves[67]
 NT 


Unknown  [67]

Indochinese mouse-eared bat


M. indochinensis
Son, Motokawa, Estók, Thong, Dang, Oshida, Csorba, Francis, Görföl, & Endō, 2013
Vietnam and China
 
Size: Unknown length
4–5 cm (2–2 in) arm/wing length[10]

Habitat: Forest[68]
 DD 


Unknown  [68]

Insular myotis


M. insularum
Dobson, 1878
American Samoa Size: About 4 cm (2 in), plus about 4 cm (2 in) tail
about 4 cm (2 in) arm/wing length[10]

Habitat: Unknown[69]
 DD 


Unknown  [69]

Izecksohn's myotis


M. izecksohni
Moratelli, Peracchi, Dias, & de Oliveira, 2011
Brazil and Argentina
 
Size: 4–7 cm (2–3 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest[70]
 DD 


Unknown  [70]

Kashmir cave bat


M. longipes
Dobson, 1873
Southern Asia Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[10]

Habitat: Forest and caves[71]
 DD 


Unknown  [71]

Keen's myotis

 

M. keenii
(Merriam, 1895)
Western Canada and United States
 
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Forest and rocky areas[72]
 LC 


Unknown  [72]

Kei myotis


M. stalkeri
Thomas, 1910
Indonesia
 
Size: 4–8 cm (2–3 in), plus 5–6 cm (2–2 in) tail
5–6 cm (2–2 in) arm/wing length[10]

Habitat: Forest, inland wetlands, and caves[73]
 LC 


10,000–11,000  [73]

Kock's mouse-eared bat


M. dieteri
Happold, 2005
Republic of the Congo Size: About 5 cm (2 in), plus about 4 cm (2 in) tail
about 4 cm (2 in) arm/wing length[7]

Habitat: Forest and caves[74]
 DD 


Unknown  [74]

Large myotis


M. chinensis
Tomes, 1857
Eastern and southeastern Asia
 
Size: 9–10 cm (4–4 in), plus 5–6 cm (2–2 in) tail
6–7 cm (2–3 in) arm/wing length[18]

Habitat: Forest, inland wetlands, and caves[75]
 LC 


Unknown  [75]

Large-footed bat

 

M. adversus
Horsfield, 1824

Six subspecies
  • M. a. adversus
  • M. a. carimatae
  • M. a. orientis
  • M. a. taiwanensis
  • M. a. tanimbarensis
  • M. a. wetarensis
Southeastern Asia
 
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[10]

Habitat: Forest, inland wetlands, and caves[76]
 LC 


Unknown  [76]

Large-footed myotis

 

M. macropus
(Gould, 1854)
Eastern Australia Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Inland wetlands and caves[77]
 LC 


Unknown  [77]

LaVal's myotis


M. lavali
Moratelli, Peracchi, Dias, & de Oliveira, 2011
Central and eastern South America
 
Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest, savanna, and grassland[78]
 LC 


Unknown  [78]

Lesser large-footed bat


M. hasseltii
Temminck, 1840

Four subspecies
  • M. h. abboti
  • M. h. continentis
  • M. h. hasseltii
  • M. h. macellus
Southern and southeastern Asia
 
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest and caves[79]
 LC 


Unknown  [79]

Lesser mouse-eared bat

 

M. blythii
Tomes, 1857

Four subspecies
  • M. b. ancilla
  • M. b. blythii
  • M. b. lesviacus
  • M. b. omari
Europe and Asia
 
Size: 5–8 cm (2–3 in), plus about 6 cm (2 in) tail
5–7 cm (2–3 in) arm/wing length[18]

Habitat: Shrubland, grassland, and caves[80]
 LC 


Unknown  [80]

Little brown bat

 

M. lucifugus
(Conte, 1831)

Five subspecies
  • M. l. alascensis
  • M. l. carissima
  • M. l. lucifugus
  • M. l. pernox
  • M. l. relictus
United States and Canada
 
Size: 3–6 cm (1–2 in), plus 2–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[5]

Habitat: Forest and caves[81]
 EN 


Unknown  [81]

Long-eared myotis

 

M. evotis
(H. Allen, 1864)

Six subspecies
  • M. e. chrysonotus
  • M. e. evotis
  • M. e. jonesorum
  • M. e. micronyx
  • M. e. milleri
  • M. e. pacificus
Western North America
 
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Caves, rocky areas, and forest[82]
 LC 


Unknown  [82]

Long-fingered bat

 

M. capaccinii
Bonaparte, 1837
Southern Europe, northern Africa, and western Asia
 
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[10]

Habitat: Shrubland, inland wetlands, and caves[83]
 VU 


Unknown  [83]

Long-legged myotis

 

M. volans
H. Allen, 1866

Four subspecies
  • M. v. amotus
  • M. v. interior
  • M. v. longicrus
  • M. v. volans
Western North America
 
Size: 4–7 cm (2–3 in), plus 4–5 cm (2–2 in) tail
3–5 cm (1–2 in) arm/wing length[5]

Habitat: Forest, rocky areas, caves, and desert[84]
 LC 


Unknown  [84]

Long-tailed myotis


M. longicaudatus
Ogniov, 1927

Four subspecies
  • M. l. eniseensis
  • M. l. kaguyae
  • M. l. longicaudatus
Eastern Asia (excluding red)
 
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest, inland wetlands, and caves[85]
 LC 


Unknown  [85]

Long-toed myotis

 

M. secundus
Ruedi, Csorba, Lin, & Chou, 2015
Taiwan Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[10]

Habitat: Forest[86]
 LC 


Unknown  [86]

Malagasy mouse-eared bat


M. goudoti
Smith, 1834
Madagascar
 
Size: Unknown length
3–4 cm (1–2 in) arm/wing length[7]

Habitat: Forest, rocky areas, and caves[87]
 LC 


Unknown  [87]

Malaysian whiskered myotis


M. federatus
Thomas, 1916
Malaysia (in blue)
 
Size: About 5 cm (2 in), plus about 4 cm (2 in) tail
3–5 cm (1–2 in) arm/wing length[10]

Habitat: Forest[88]
 DD 


Unknown  [88]

Maluku myotis


M. moluccarum
Thomas, 1915

Three subspecies
  • M. m. moluccarum
  • M. m. richardsi
  • M. m. solomonis
Indonesia
 
Size: 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[10]

Habitat: Forest and inland wetlands[89]
 LC 


Unknown  [89]

Mandelli's mouse-eared bat


M. sicarius
Thomas, 1915
India and Nepal Size: 5–6 cm (2–2 in), plus 4–6 cm (2–2 in) tail
4–6 cm (2–2 in) arm/wing length[18]

Habitat: Forest and caves[90]
 VU 


Unknown  [90]

Montane myotis


M. oxyotus
Peters, 1867

Two subspecies
  • M. o. gardneri
  • M. o. oxyotus
Northern and western South America
 
Size: 4–6 cm (2–2 in), plus 4–5 cm (2–2 in) tail
3–5 cm (1–2 in) arm/wing length[14]

Habitat: Forest[91]
 LC 


Unknown  [91]

Morris's bat


M. morrisi
Hill, 1971
Ethiopia and Nigeria
 
Size: About 5 cm (2 in), plus about 5 cm (2 in) tail
4–5 cm (2–2 in) arm/wing length[7]

Habitat: Savanna and caves[92]
 DD 


Unknown  [92]

Natterer's bat

 

M. nattereri
(Kuhl, 1817)

Two subspecies
  • M. n. nattereri
  • M. n. tschuliensis
Europe and western Asia Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest, shrubland, grassland, inland wetlands, and caves[93]
 LC 


Unknown  [93]

Nepal myotis


M. nipalensis
Dobson, 1871

Three subspecies
  • M. n. nipalensis
  • M. n. przewalskii
  • M. n. transcaspicus
Central and western Asia
 
Size: About 4 cm (2 in), plus about 33–5 cm (1–2 in) tail
about 3 cm (1 in) arm/wing length[7]

Habitat: Forest, shrubland, grassland, caves, and desert[94]
 LC 


Unknown  [94]

Nimba myotis


M. nimbaensis
Simmons, Flanders, Fils, Parker, Suter, Bamba, Douno, Keita, Morales, & Frick, 2021
Guinea Size: 6–8 cm (2–3 in), plus 4–5 cm (2–2 in) tail
5–6 cm (2–2 in) arm/wing length[7]

Habitat: Forest, grassland, inland wetlands, caves, and shrubland[95]
 CR 


Unknown  [95]

Northern long-eared bat

 

M. septentrionalis
Trouessart, 1897
Canada and eastern United States
 
Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Forest and caves[96]
 NT 


Unknown  [96]

Orange-fingered myotis


M. rufopictus
(Waterhouse, 1845)
Indonesia
 
Size: About 6 cm (2 in), plus 4–6 cm (2–2 in) tail
4–6 cm (2–2 in) arm/wing length[7]

Habitat: Forest[97]
 DD 


Unknown  [97]

Pallid large-footed myotis


M. macrotarsus
Waterhouse, 1845

Two subspecies
  • M. m. macrotarsus
  • M. m. saba
Indonesia
 
Size: About 6 cm (2 in), plus 4–6 cm (2–2 in) tail
4–5 cm (2–2 in) arm/wing length[10]

Habitat: Caves[98]
 LC 


Unknown  [98]

Peninsular myotis

 

M. peninsularis
Miller, 1898
Southern Baja California peninsula
 
Size: 4–6 cm (2–2 in), plus 3–6 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[14]

Habitat: Forest and caves[99]
 EN 


Unknown  [99]

Peters's myotis


M. ater
(Peters, 1866)

Two subspecies
  • M. a. ater
  • M. a. nugax
Southeastern Asia
 
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[10]

Habitat: Forest[100]
 LC 


Unknown  [100]

Peyton's myotis


M. peytoni
Wroughton & Ryley, 1913
Southern India (in yellow)
 
Size: 5–7 cm (2–3 in), plus 4–5 cm (2–2 in) tail
4–5 cm (2–2 in) arm/wing length[10]

Habitat: Forest[101]
 DD 


Unknown  [101]

Pond bat

 

M. dasycneme
(Boie, 1825)
Europe and northern Asia
 
Size: 5–7 cm (2–3 in), plus 4–6 cm (2–2 in) tail
4–5 cm (2–2 in) arm/wing length[7]

Habitat: Forest, inland wetlands, and caves[102]
 NT 


Unknown  [102]

Red myotis

 

M. ruber
Geoffroy, 1806
Southeastern South America
 
Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[5]

Habitat: Forest[103]
 NT 


Unknown  [103]

Reddish myotis


M. soror
Ruedi, Csorba, Lin, & Chou, 2015
Taiwan Size: About 5 cm (2 in), plus about 4 cm (2 in) tail
about 4 cm (2 in) arm/wing length[18]

Habitat: Forest[104]
 DD 


Unknown  [104]

Reddish-black myotis

 

M. rufoniger
(Tomes, 1858)
Eastern Asia
 
Size: 5–8 cm (2–3 in), plus 4–6 cm (2–2 in) tail
4–6 cm (2–2 in) arm/wing length[7]

Habitat: Forest and caves[105]
 LC 


Unknown  [105]

Rickett's big-footed bat


M. pilosus
Peters, 1869
Eastern Asia
 
Size: 5–7 cm (2–3 in), plus 3–6 cm (1–2 in) tail
5–7 cm (2–3 in) arm/wing length[18]

Habitat: Forest and inland wetlands[106]
 VU 


Unknown  [106]

Ridley's bat


M. ridleyi
Thomas, 1898
Southeastern Asia
 
Size: 3–5 cm (1–2 in), plus 2–4 cm (1–2 in) tail
2–4 cm (1–2 in) arm/wing length[10]

Habitat: Caves, inland wetlands, and forest[107]
 NT 


Unknown  [107]

Riparian myotis


M. riparius
Handley, 1960
Central America and South America
 
Size: 4–5 cm (2–2 in), plus 2–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest[108]
 LC 


Unknown  [108]

Rufous mouse-eared bat

 

M. bocagii
Peters, 1870

Three subspecies
  • M. b. bocagii
  • M. b. cupreolus
  • M. b. dogalensis
Scattered sub-Saharan Africa and Yemen
 
Size: 5–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[7]

Habitat: Forest and savanna[109]
 LC 


Unknown  [109]

Schaub's myotis


M. schaubi
Kormos, 1934

Two subspecies
  • M. s. araxenus
  • M. s. schaubi
Armenia and Iran Size: 4–6 cm (2–2 in), plus 4–5 cm (2–2 in) tail
3–5 cm (1–2 in) arm/wing length[18]

Habitat: Forest, shrubland, and caves[110]
 DD 


Unknown  [110]

Schwartz's myotis


M. martiniquensis
LaVal, 1973
Barbados and Martinique in the Caribbean
 
Size: Unknown length
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Caves[111]
 NT 


Unknown  [111]

Scott's mouse-eared bat


M. scotti
Thomas, 1927
Ethiopia Size: 4–6 cm (2–2 in), plus 4–5 cm (2–2 in) tail
3–5 cm (1–2 in) arm/wing length[7]

Habitat: Forest and shrubland[112]
 VU 


Unknown  [112]

Siberian bat

 

M. sibiricus
(Kastschenko, 1905)
Northeastern Asia (in green)
 
Size: 3–6 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[7]

Habitat: Forest, savanna, rocky areas, and caves[113]
 LC 


Unknown  [113]

Silver-tipped myotis

 

M. albescens
Geoffroy, 1806
Central America and South America
 
Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest[114]
 LC 


Unknown  [114]

Singapore whiskered bat


M. oreias
(Temminck, 1840)
Singapore
 
Size: Unknown[115]

Habitat: Unknown[115]
 DD 


Unknown  [115]

Southeastern myotis

 

M. austroriparius
(Rhoads, 1897)
Southeastern United States
 
Size: 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest and caves[116]
 LC 


Unknown  [116]

Southern myotis


M. aelleni
Baud, 1979
Argentina
 
Size: 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length[14]

Habitat: Unknown[117]
 DD 


Unknown  [117]

Southwestern myotis

 

M. auriculus
Baker & Stains, 1955

Two subspecies
  • M. a. apache
  • M. a. auriculus
Southern United States and Mexico
 
Size: 5–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Forest, caves, and desert[118]
 LC 


Unknown  [118]

Szechwan myotis


M. altarium
Thomas, 1911
China and Thailand
 
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
4–5 cm (2–2 in) arm/wing length[7]

Habitat: Caves[119]
 LC 


Unknown  [119]

Thick-thumbed myotis


M. rosseti
Oei, 1951
Southeastern Asia
 
Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail
2–4 cm (1–2 in) arm/wing length[10]

Habitat: Forest[120]
 LC 


Unknown  [120]

Velvety myotis


M. simus
Thomas, 1901
Western, central, and eastern South America
 
Size: 5–6 cm (2–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest[121]
 DD 


Unknown  [121]

Wall-roosting mouse-eared bat

 

M. muricola
(Gray, 1846)

Seven subspecies
  • M. m. browni
  • M. m. caliginosus
  • M. m. herrei
  • M. m. moupinensis
  • M. m. muricola
  • M. m. niasensis
  • M. m. patriciae
Southeastern Asia
 
Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[10]

Habitat: Forest and caves[122]
 LC 


Unknown  [122]

Weber's myotis


M. weberi
(Jentink, 1890)
Sulawesi island in Indonesia Size: About 6 cm (2 in), plus about 4 cm (2 in) tail
4–6 cm (2–2 in) arm/wing length[7]

Habitat: Forest[123]
 DD 


Unknown  [123]

Welwitsch's bat

 

M. welwitschii
(Gray, 1866)
Western, eastern, and southern Africa
 
Size: 5–7 cm (2–3 in), plus 5–7 cm (2–3 in) tail
5–6 cm (2–2 in) arm/wing length[7]

Habitat: Forest, savanna, shrubland, and grassland[124]
 LC 


Unknown  [124]

Western small-footed bat

 

M. ciliolabrum
(Merriam, 1842)
Western North America
 
Size: 3–5 cm (1–2 in), plus 2–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Forest, rocky areas, caves, and desert[125]
 LC 


Unknown  [125]

Whiskered bat

 

M. mystacinus
(Kuhl, 1817)

Three subspecies
  • M. m. caucasicus
  • M. m. mystacinus
  • M. m. occidentalis
Europe, northern Africa, and western Asia
 
Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[7]

Habitat: Desert, caves, grassland, shrubland, and forest[126]
 LC 


Unknown  [126]

Yanbaru whiskered bat


M. yanbarensis
Maeda & Matsumura, 1998
Ryukyu Islands in Japan
 
Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[10]

Habitat: Forest[127]
 CR 


Unknown  [127]

Yellowish myotis

 

M. levis
Geoffroy, 1806
Southeastern South America (in red)
 
Size: 3–6 cm (1–2 in), plus 4–5 cm (2–2 in) tail
3–5 cm (1–2 in) arm/wing length[14]

Habitat: Forest[128]
 LC 


Unknown  [128]

Yuma myotis

 

M. yumanensis
H. Allen, 1864

Six subspecies
  • M. y. lambi
  • M. y. lutosus
  • M. y. oxalis
  • M. y. saturatus
  • M. y. sociabilis
  • M. y. yumanensis
Western North America
 
Size: 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[14]

Habitat: Forest and caves[129]
 LC 


Unknown  [129]

Genus Submyotodon Ziegler, 2003 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Taiwan broad-muzzled bat


S. latirostris
(Kishida, 1932)
Taiwan Size: 3–4 cm (1–2 in), plus 3–4 cm (1–2 in) tail
3–4 cm (1–2 in) arm/wing length[5]

Habitat: Forest[130]
 LC 


Unknown  [130]

References

edit
  1. ^ Nowak, pp. 184–186
  2. ^ Levin, E.; Barnea, A.; Yovel, Y.; Yom-Tov, Y. (2006). "Have introduced fish initiated piscivory among the long-fingered bat?". Mammalian Biology. 71 (3): 139–143. Bibcode:2006MamBi..71..139L. doi:10.1016/j.mambio.2006.01.002.
  3. ^ "Fossilworks: Myotis". Paleobiology Database. University of Wisconsin–Madison. Archived from the original on September 22, 2022. Retrieved November 2, 2024.
  4. ^ Wilson, Reeder, pp. 499–518
  5. ^ a b c d e f g h i j k l m n o p q r s t Chernasky; Motis; Burgin, pp. 556–557
  6. ^ a b Soisook, P.; Csorba, G.; Bumrungsri, S.; Francis, C. M.; Bates, P.; Kingston, T. (2016). "Eudiscopus denticulus". IUCN Red List of Threatened Species. 2016: e.T8168A22028419. doi:10.2305/IUCN.UK.2016-2.RLTS.T8168A22028419.en.
  7. ^ a b c d e f g h i j k l m n o p q r s t u v w x y Chernasky; Motis; Burgin, pp. 560–561
  8. ^ a b Hutson, A. M.; Paunović, M. (2016). "Myotis alcathoe". IUCN Red List of Threatened Species. 2016: e.T136680A518740. doi:10.2305/IUCN.UK.2016-2.RLTS.T136680A518740.en.
  9. ^ a b Jacobs, D. (2019). "Myotis anjouanensis". IUCN Red List of Threatened Species. 2019: e.T44863A22073545. doi:10.2305/IUCN.UK.2019-3.RLTS.T44863A22073545.en.
  10. ^ a b c d e f g h i j k l m n o p q r s t u v w x y z Chernasky; Motis; Burgin, pp. 562–563
  11. ^ a b Srinivasulu, C.; Srinivasulu, B. (2019). "Myotis annatessae". IUCN Red List of Threatened Species. 2019: e.T85342605A85342608. doi:10.2305/IUCN.UK.2019-3.RLTS.T85342605A85342608.en.
  12. ^ a b Kruskop, S. V. (2016). "Myotis annamiticus". IUCN Red List of Threatened Species. 2016: e.T136279A22006224. doi:10.2305/IUCN.UK.2016-2.RLTS.T136279A22006224.en.
  13. ^ a b Solari, S. (2018). "Myotis occultus". IUCN Red List of Threatened Species. 2018: e.T136650A21990499. doi:10.2305/IUCN.UK.2018-2.RLTS.T136650A21990499.en.
  14. ^ a b c d e f g h i j k l m n o p q r s t u v w x Chernasky; Motis; Burgin, pp. 558–559
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  17. ^ a b Larsen, R. (2016). "Myotis nyctor". IUCN Red List of Threatened Species. 2016: e.T76435059A76435083. doi:10.2305/IUCN.UK.2016-1.RLTS.T76435059A76435083.en.
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  24. ^ a b Gazaryan, S.; Kruskop, S. V.; Godlevska, L. (2021) [errata version of 2020 assessment]. "Myotis brandtii". IUCN Red List of Threatened Species. 2020: e.T85566997A195857637. doi:10.2305/IUCN.UK.2020-2.RLTS.T85566997A195857637.en.
  25. ^ a b Görföl, T.; Tu, V.; Csorba, G.; Francis, C. M.; Hutson, A. M.; Bates, P. J. J.; Bumrungsri, S.; Molur, S.; Srinivasulu, C. (2020). "Myotis montivagus". IUCN Red List of Threatened Species. 2020: e.T85567622A22065126. doi:10.2305/IUCN.UK.2020-2.RLTS.T85567622A22065126.en.
  26. ^ a b Arroyo-Cabrales, J.; Perez, S. (2017). "Myotis californicus". IUCN Red List of Threatened Species. 2017: e.T14150A22061366. doi:10.2305/IUCN.UK.2017-2.RLTS.T14150A22061366.en.
  27. ^ a b Monadjem, A.; Jacobs, D. (2017). "Myotis tricolor". IUCN Red List of Threatened Species. 2017: e.T14207A22063832. doi:10.2305/IUCN.UK.2017-2.RLTS.T14207A22063832.en.
  28. ^ a b Solari, S. (2019). "Myotis velifer". IUCN Red List of Threatened Species. 2019: e.T14208A22063586. doi:10.2305/IUCN.UK.2019-1.RLTS.T14208A22063586.en.
  29. ^ a b Barquez, R.; Diaz, M. (2016). "Myotis chiloensis". IUCN Red List of Threatened Species. 2016: e.T14151A22061103. doi:10.2305/IUCN.UK.2016-3.RLTS.T14151A22061103.en.
  30. ^ a b Feng, J.; Jiang, T. L. (2019). "Myotis laniger". IUCN Red List of Threatened Species. 2019: e.T136429A21984685. doi:10.2305/IUCN.UK.2019-3.RLTS.T136429A21984685.en.
  31. ^ a b Perez, S.; de Grammont, P. C.; Cuarón, A. D. (2017). "Myotis fortidens". IUCN Red List of Threatened Species. 2017: e.T14161A22056846. doi:10.2305/IUCN.UK.2017-2.RLTS.T14161A22056846.en.
  32. ^ a b Russo, D.; Cistrone, L. (2024) [errata version of 2023 assessment]. "Myotis crypticus". IUCN Red List of Threatened Species. 2023: e.T215154989A254355251.
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  34. ^ a b Solari, S. (2016). "Myotis nesopolus". IUCN Red List of Threatened Species. 2016: e.T14184A22065759. doi:10.2305/IUCN.UK.2016-1.RLTS.T14184A22065759.en.
  35. ^ a b Arroyo-Cabrales, J.; Álvarez-Castañeda, S. T. (2017). "Myotis melanorhinus". IUCN Red List of Threatened Species. 2017: e.T136784A22033542. doi:10.2305/IUCN.UK.2017-2.RLTS.T136784A22033542.en.
  36. ^ a b Kruskop, S. V.; Godlevska, L.; Bücs, S.; Çoraman, E.; Gazaryan, S. (2021) [errata version of 2020 assessment]. "Myotis daubentonii". IUCN Red List of Threatened Species. 2020: e.T85342710A195858793. doi:10.2305/IUCN.UK.2020-2.RLTS.T85342710A195858793.en.
  37. ^ a b Jiang, T. L.; Feng, J. (2019). "Myotis davidii". IUCN Red List of Threatened Species. 2019: e.T136250A22003049. doi:10.2305/IUCN.UK.2019-3.RLTS.T136250A22003049.en.
  38. ^ a b Solari, S. (2017). "Myotis diminutus". IUCN Red List of Threatened Species. 2017: e.T88151417A88151431. doi:10.2305/IUCN.UK.2017-2.RLTS.T88151417A88151431.en.
  39. ^ a b Barquez, R.; Diaz, M. (2016). "Myotis dinellii". IUCN Red List of Threatened Species. 2016: e.T136204A22009702. doi:10.2305/IUCN.UK.2016-3.RLTS.T136204A22009702.en.
  40. ^ a b Larsen, R. (2016). "Myotis dominicensis". IUCN Red List of Threatened Species. 2016: e.T14155A22057933. doi:10.2305/IUCN.UK.2016-1.RLTS.T14155A22057933.en.
  41. ^ a b Fukui, D.; Sun, K.; Kruskop, S. V. (2019). "Myotis macrodactylus". IUCN Red List of Threatened Species. 2019: e.T14177A22065868. doi:10.2305/IUCN.UK.2019-3.RLTS.T14177A22065868.en.
  42. ^ a b Solari, S. (2018). "Myotis leibii". IUCN Red List of Threatened Species. 2018: e.T14172A22055716. doi:10.2305/IUCN.UK.2018-2.RLTS.T14172A22055716.en.
  43. ^ a b Fukui, D.; Sano, A. (2020). "Myotis petax". IUCN Red List of Threatened Species. 2020: e.T85342726A85342734. doi:10.2305/IUCN.UK.2020-2.RLTS.T85342726A85342734.en.
  44. ^ a b Miller, B.; Rodriguez, B. (2017) [errata version of 2016 assessment]. "Myotis elegans". IUCN Red List of Threatened Species. 2016: e.T14156A115121563. doi:10.2305/IUCN.UK.2016-3.RLTS.T14156A22057814.en.
  45. ^ a b Russo, D.; Cistrone, L. (2023). "Myotis escalerai". IUCN Red List of Threatened Species. 2023: e.T85733126A211003991. doi:10.2305/IUCN.UK.2023-1.RLTS.T85733126A211003991.en.
  46. ^ a b Fukui, D.; Sano, A.; Kruskop, S. V. (2019). "Myotis bombinus". IUCN Red List of Threatened Species. 2019: e.T14149A22061650. doi:10.2305/IUCN.UK.2019-3.RLTS.T14149A22061650.en.
  47. ^ a b Juste, J.; Paunović, M. (2016). "Myotis punicus". IUCN Red List of Threatened Species. 2016: e.T44864A22073410. doi:10.2305/IUCN.UK.2016-2.RLTS.T44864A22073410.en.
  48. ^ a b Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis vivesi". IUCN Red List of Threatened Species. 2016: e.T14209A22069146. doi:10.2305/IUCN.UK.2016-1.RLTS.T14209A22069146.en.
  49. ^ a b Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis findleyi". IUCN Red List of Threatened Species. 2016: e.T14159A22058800. doi:10.2305/IUCN.UK.2016-1.RLTS.T14159A22058800.en.
  50. ^ a b Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis planiceps". IUCN Red List of Threatened Species. 2016: e.T14191A22066742. doi:10.2305/IUCN.UK.2016-1.RLTS.T14191A22066742.en.
  51. ^ a b Vincenot, C. E.; Preble, J. H.; Huang, J. C. -C.; Collazo, A. M.; Kamal, A. (2021). "Myotis frater". IUCN Red List of Threatened Species. 2021: e.T85566806A22056940. doi:10.2305/IUCN.UK.2021-2.RLTS.T85566806A22056940.en.
  52. ^ a b Jiang, T. L.; Feng, J. (2019). "Myotis fimbriatus". IUCN Red List of Threatened Species. 2019: e.T85735587A22058886. doi:10.2305/IUCN.UK.2019-3.RLTS.T85735587A22058886.en.
  53. ^ a b Arroyo-Cabrales, J.; de Grammont, P. C. (2017). "Myotis thysanodes". IUCN Red List of Threatened Species. 2017: e.T14206A22063246. doi:10.2305/IUCN.UK.2017-2.RLTS.T14206A22063246.en.
  54. ^ a b Fukui, D.; Sano, A. (2021) [errata version of 2019 assessment]. "Myotis pruinosus". IUCN Red List of Threatened Species. 2019: e.T14192A209551299. doi:10.2305/IUCN.UK.2019-3.RLTS.T14192A209551299.en.
  55. ^ a b Piraccini, R. (2016). "Myotis emarginatus". IUCN Red List of Threatened Species. 2016: e.T14129A22051191. doi:10.2305/IUCN.UK.2016-2.RLTS.T14129A22051191.en.
  56. ^ a b Waldien, D. L.; Santiago, K.; Wortham, G.; Stronsick, S. (2021). "Myotis gomantongensis". IUCN Red List of Threatened Species. 2021: e.T40035A22060096. doi:10.2305/IUCN.UK.2021-3.RLTS.T40035A22060096.en.
  57. ^ a b Solari, S. (2018). "Myotis grisescens". IUCN Red List of Threatened Species. 2018: e.T14132A22051652. doi:10.2305/IUCN.UK.2018-2.RLTS.T14132A22051652.en.
  58. ^ a b Coroiu, I.; Juste, J.; Paunović, M. (2016). "Myotis myotis". IUCN Red List of Threatened Species. 2016: e.T14133A22051759. doi:10.2305/IUCN.UK.2016-2.RLTS.T14133A22051759.en.
  59. ^ a b Cajas C., J.; Miller, B. (2016). "Myotis cobanensis". IUCN Red List of Threatened Species. 2016: e.T14154A22058031. doi:10.2305/IUCN.UK.2016-2.RLTS.T14154A22058031.en.
  60. ^ a b Görföl, T.; Csorba, G.; Bates, P. J. J.; Francis, C. M.; Molur, S.; Srinivasulu, C. (2020). "Myotis annectans". IUCN Red List of Threatened Species. 2020: e.T14142A22050272. doi:10.2305/IUCN.UK.2020-2.RLTS.T14142A22050272.en.
  61. ^ a b Barquez, R.; Diaz, M. (2016). "Myotis keaysi". IUCN Red List of Threatened Species. 2016: e.T14170A22056048. doi:10.2305/IUCN.UK.2016-3.RLTS.T14170A22056048.en.
  62. ^ a b Csorba, G.; Hutson, A. M.; Kingston, T.; Bumrungsri, S.; Francis, C. M. (2016). "Myotis hermani". IUCN Red List of Threatened Species. 2016: e.T14165A22057251. doi:10.2305/IUCN.UK.2016-2.RLTS.T14165A22057251.en.
  63. ^ a b Santiago, K.; Wortham, G.; Waldien, D. L. (2021). "Myotis siligorensis". IUCN Red List of Threatened Species. 2021: e.T14203A22064839. doi:10.2305/IUCN.UK.2021-3.RLTS.T14203A22064839.en.
  64. ^ a b Huang, J. C. -C.; Csorba, G.; Chang, H.-C.; Ho, Y.-Y. (2020). "Myotis formosus". IUCN Red List of Threatened Species. 2020: e.T85736120A95642290. doi:10.2305/IUCN.UK.2020-2.RLTS.T85736120A95642290.en.
  65. ^ a b Phelps, K.; Rosell-Ambal, R. G. B.; Tabaranza, B.; Heaney, L.; Gonzalez, J. C.; Molur, S.; Srinivasulu, C. (2019). "Myotis horsfieldii". IUCN Red List of Threatened Species. 2019: e.T14166A22057415. doi:10.2305/IUCN.UK.2019-3.RLTS.T14166A22057415.en.
  66. ^ a b Zhigalin, A.; Stubbe, M.; Ariunbold, J.; Buuveibaatar, V.; Dorjderem, S.; Monkhzul, T.; Otgonbaatar, M.; Tsogbadrakh, M. (2020). "Myotis ikonnikovi". IUCN Red List of Threatened Species. 2020: e.T14168A22057122. doi:10.2305/IUCN.UK.2020-3.RLTS.T14168A22057122.en.
  67. ^ a b Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis sodalis". IUCN Red List of Threatened Species. 2016: e.T14136A22053184. doi:10.2305/IUCN.UK.2016-1.RLTS.T14136A22053184.en.
  68. ^ a b Son, N.; Görföl, T.; Csorba, G. (2019). "Myotis indochinensis". IUCN Red List of Threatened Species. 2019: e.T85342688A85342691. doi:10.2305/IUCN.UK.2019-3.RLTS.T85342688A85342691.en.
  69. ^ a b Helgen, K.; Bonaccorso, F. J. (2020). "Myotis insularum". IUCN Red List of Threatened Species. 2020: e.T14169A22055968. doi:10.2305/IUCN.UK.2020-2.RLTS.T14169A22055968.en.
  70. ^ a b Solari, S. (2017). "Myotis izecksohni". IUCN Red List of Threatened Species. 2017: e.T88151563A88151572. doi:10.2305/IUCN.UK.2017-2.RLTS.T88151563A88151572.en.
  71. ^ a b Kruskop, S. V. (2016). "Myotis longipes". IUCN Red List of Threatened Species. 2016: e.T14175A22056206. doi:10.2305/IUCN.UK.2016-2.RLTS.T14175A22056206.en.
  72. ^ a b Arroyo-Cabrales, J.; Álvarez-Castañeda, S. T. (2017). "Myotis keenii". IUCN Red List of Threatened Species. 2017: e.T14171A22055579. doi:10.2305/IUCN.UK.2017-2.RLTS.T14171A22055579.en.
  73. ^ a b Bouillard, N. (2021). "Myotis stalkeri". IUCN Red List of Threatened Species. 2021: e.T14205A22063416. doi:10.2305/IUCN.UK.2021-3.RLTS.T14205A22063416.en.
  74. ^ a b Happold. M. (2019). "Myotis dieteri". IUCN Red List of Threatened Species. 2019: e.T136678A22038629. doi:10.2305/IUCN.UK.2019-3.RLTS.T136678A22038629.en.
  75. ^ a b Jiang, T. L.; Feng, J. (2019). "Myotis chinensis". IUCN Red List of Threatened Species. 2019: e.T14152A22060946. doi:10.2305/IUCN.UK.2019-3.RLTS.T14152A22060946.en.
  76. ^ a b Bouillard, N. (2021). "Myotis adversus". IUCN Red List of Threatened Species. 2021: e.T85735326A22049231. doi:10.2305/IUCN.UK.2021-2.RLTS.T85735326A22049231.en.
  77. ^ a b Gorecki, V.; Pennay, M. (2021). "Myotis macropus". IUCN Red List of Threatened Species. 2021: e.T136697A22039960. doi:10.2305/IUCN.UK.2021-1.RLTS.T136697A22039960.en.
  78. ^ a b Solari, S. (2017). "Myotis lavali". IUCN Red List of Threatened Species. 2017: e.T88151601A88151604. doi:10.2305/IUCN.UK.2017-2.RLTS.T88151601A88151604.en.
  79. ^ a b Bates, P. J. J.; Hutson, A. M.; Carino, A.; Kingston, T.; Lee, B. P. Y-H; Maryanto, I.; Molur, S.; Srinivasulu, C. (2020). "Myotis hasseltii". IUCN Red List of Threatened Species. 2020: e.T14164A22056644. doi:10.2305/IUCN.UK.2020-2.RLTS.T14164A22056644.en.
  80. ^ a b Juste, J.; Paunović, M. (2016). "Myotis blythii". IUCN Red List of Threatened Species. 2016: e.T14124A22053297. doi:10.2305/IUCN.UK.2016-2.RLTS.T14124A22053297.en.
  81. ^ a b Solari, S. (2021) [amended version of 2018 assessment]. "Myotis lucifugus". IUCN Red List of Threatened Species. 2021: e.T14176A208031565. doi:10.2305/IUCN.UK.2021-3.RLTS.T14176A208031565.en.
  82. ^ a b Arroyo-Cabrales, J.; Álvarez-Castañeda, S. T. (2017). "Myotis evotis". IUCN Red List of Threatened Species. 2017: e.T14157A22059133. doi:10.2305/IUCN.UK.2017-2.RLTS.T14157A22059133.en.
  83. ^ a b Paunović, M. (2016). "Myotis capaccinii". IUCN Red List of Threatened Species. 2016: e.T14126A22054131. doi:10.2305/IUCN.UK.2016-2.RLTS.T14126A22054131.en.
  84. ^ a b Solari, S. (2019). "Myotis volans". IUCN Red List of Threatened Species. 2019: e.T14210A22069325. doi:10.2305/IUCN.UK.2019-1.RLTS.T14210A22069325.en.
  85. ^ a b Vincenot, C. E.; Preble, J. H.; Collazo, A. M. (2021). "Myotis longicaudatus". IUCN Red List of Threatened Species. 2021: e.T85566977A85566980. doi:10.2305/IUCN.UK.2021-2.RLTS.T85566977A85566980.en.
  86. ^ a b Ruedi, M.; Csorba, G.; Liang-Kong, L.; Cheng-Han, C. (2017). "Myotis secundus". IUCN Red List of Threatened Species. 2017: e.T85342651A85342654. doi:10.2305/IUCN.UK.2017-2.RLTS.T85342651A85342654.en.
  87. ^ a b Monadjem, A.; Razafimanahaka, J.; Ranivo, J.; Kofoky, A.; Hutson, A. M.; Cardiff, S. G.; Andriafidison, D.; Goodman, S.; Jenkins, R. K. B.; Racey, P. A.; Ratrimomanarivo, F. H. (2017). "Myotis goudoti". IUCN Red List of Threatened Species. 2017: e.T14163A22056541. doi:10.2305/IUCN.UK.2017-2.RLTS.T14163A22056541.en.
  88. ^ a b Görföl, T.; Csorba, G. (2017). "Myotis federatus". IUCN Red List of Threatened Species. 2017: e.T85568302A85568305. doi:10.2305/IUCN.UK.2017-2.RLTS.T85568302A85568305.en.
  89. ^ a b Armstrong, K. N. (2021). "Myotis moluccarum". IUCN Red List of Threatened Species. 2021: e.T136770A22033795. doi:10.2305/IUCN.UK.2021-3.RLTS.T136770A22033795.en.
  90. ^ a b Srinivasulu, B.; Srinivasulu, C. (2019). "Myotis sicarius". IUCN Red List of Threatened Species. 2019: e.T14202A22063965. doi:10.2305/IUCN.UK.2019-3.RLTS.T14202A22063965.en.
  91. ^ a b Solari, S. (2018). "Myotis oxyotus". IUCN Red List of Threatened Species. 2018: e.T14187A22067211. doi:10.2305/IUCN.UK.2018-2.RLTS.T14187A22067211.en.
  92. ^ a b Jacobs, D.; Cotterill, F. P. D.; Taylor, P. J. (2019). "Myotis morrisi". IUCN Red List of Threatened Species. 2019: e.T14182A22065314. doi:10.2305/IUCN.UK.2019-3.RLTS.T14182A22065314.en.
  93. ^ a b Russo, D.; Cistrone, L. (2023). "Myotis nattereri". IUCN Red List of Threatened Species. 2023: e.T215492021A211005466. doi:10.2305/IUCN.UK.2023-1.RLTS.T215492021A211005466.en.
  94. ^ a b Srinivasulu, B.; Srinivasulu, C. (2019). "Myotis nipalensis". IUCN Red List of Threatened Species. 2019: e.T136495A21976309. doi:10.2305/IUCN.UK.2019-3.RLTS.T136495A21976309.en.
  95. ^ a b Bakwo Fils, E. M.; Flanders, J.; Frick, W. F.; Simmons, N. (2022). "Myotis nimbaensis". IUCN Red List of Threatened Species. 2022: e.T216617275A216617367. doi:10.2305/IUCN.UK.2022-2.RLTS.T216617275A216617367.en.
  96. ^ a b Solari, S. (2018). "Myotis septentrionalis". IUCN Red List of Threatened Species. 2018: e.T14201A22064312. doi:10.2305/IUCN.UK.2018-2.RLTS.T14201A22064312.en.
  97. ^ a b Csorba, G.; Rosell-Ambal, R. G. B.; Tabaranza, B.; Sedlock, J.; Ingle, N. R.; Heaney, L.; Balete, D. S.; Ong, P. (2016). "Myotis rufopictus". IUCN Red List of Threatened Species. 2016: e.T136411A22017446. doi:10.2305/IUCN.UK.2016-2.RLTS.T136411A22017446.en.
  98. ^ a b Duya, M. R.; Alvarez, J.; Fidelino, J.; Gatan-Balbas, M.; Pedregosa, M.; Veluz, M. J.; Jakosalem, P. G.; Tanalgo, K.; Garcia, J. J. L.; Cacho, M. A. (2019). "Myotis macrotarsus". IUCN Red List of Threatened Species. 2019: e.T14178A22065997. doi:10.2305/IUCN.UK.2019-3.RLTS.T14178A22065997.en.
  99. ^ a b Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis peninsularis". IUCN Red List of Threatened Species. 2016: e.T14189A22066405. doi:10.2305/IUCN.UK.2016-1.RLTS.T14189A22066405.en.
  100. ^ a b Wiles, G.; Furey, N. (2021). "Myotis ater". IUCN Red List of Threatened Species. 2021: e.T14144A22050847. doi:10.2305/IUCN.UK.2021-2.RLTS.T14144A22050847.en.
  101. ^ a b Csorba, G.; Görföl, T. (2017). "Myotis peytoni". IUCN Red List of Threatened Species. 2017: e.T85568321A85568324. doi:10.2305/IUCN.UK.2017-2.RLTS.T85568321A85568324.en.
  102. ^ a b Piraccini, R. (2016). "Myotis dasycneme". IUCN Red List of Threatened Species. 2016: e.T14127A22055164. doi:10.2305/IUCN.UK.2016-2.RLTS.T14127A22055164.en.
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eth 2
orte 1
see 1