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The Cenozoic Portal

Introduction

The Cenozoic (/ˌsiːnəˈzoʊ.ɪk, ˌsɛn-/ SEE-nə-ZOH-ik, SEN-ə-; lit. 'new life') is Earth's current geological era, representing the last 66 million years of Earth's history. It is characterized by the dominance of insects, mammals, birds and angiosperms (flowering plants). It is the latest of three geological eras of the Phanerozoic Eon, preceded by the Mesozoic and Paleozoic. The Cenozoic started with the Cretaceous–Paleogene extinction event, when many species, including the non-avian dinosaurs, became extinct in an event attributed by most experts to the impact of a large asteroid or other celestial body, the Chicxulub impactor.

The Cenozoic is also known as the Age of Mammals because the terrestrial animals that dominated both hemispheres were mammals – the eutherians (placentals) in the Northern Hemisphere and the metatherians (marsupials, now mainly restricted to Australia and to some extent South America) in the Southern Hemisphere. The extinction of many groups allowed mammals and birds to greatly diversify so that large mammals and birds dominated life on Earth. The continents also moved into their current positions during this era.

The climate during the early Cenozoic was warmer than today, particularly during the Paleocene–Eocene Thermal Maximum. However, the Eocene to Oligocene transition and the Quaternary glaciation dried and cooled Earth. (Full article...)

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Selected article on the Cenozoic world and its legacies

Representatives of the three major extant spider groups (counterclockwise): Mesothelae, Araneomorphae and Mygalomorphae.

Spiders (order Araneae) are air-breathing arthropods that have eight limbs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species diversity among all orders of organisms. Spiders are found worldwide on every continent except Antarctica, and have become established in nearly every land habitat. As of September 2024^[update], 52,309 spider species in 134 families have been recorded by taxonomists. However, there has been debate among scientists about how families should be classified, with over 20 different classifications proposed since 1900.

Anatomically, spiders (as with all arachnids) differ from other arthropods in that the usual body segments are fused into two tagmata, the cephalothorax or prosoma, and the opisthosoma, or abdomen, and joined by a small, cylindrical pedicel. However, as there is currently neither paleontological nor embryological evidence that spiders ever had a separate thorax-like division, there exists an argument against the validity of the term cephalothorax, which means fused cephalon (head) and the thorax. Similarly, arguments can be formed against the use of the term "abdomen", as the opisthosoma of all spiders contains a heart and respiratory organs, organs atypical of an abdomen.

Unlike insects, spiders do not have antennae. In all except the most primitive group, the Mesothelae, spiders have the most centralized nervous systems of all arthropods, as all their ganglia are fused into one mass in the cephalothorax. Unlike most arthropods, spiders have no extensor muscles in their limbs and instead extend them by hydraulic pressure.

Their abdomens bear appendages, modified into spinnerets that extrude silk from up to six types of glands. Spider webs vary widely in size, shape and the amount of sticky thread used. It now appears that the spiral orb web may be one of the earliest forms, and spiders that produce tangled cobwebs are more abundant and diverse than orb-weaver spiders. Spider-like arachnids with silk-producing spigots (Uraraneida) appeared in the Devonian period, about 386 million years ago, but these animals apparently lacked spinnerets. True spiders have been found in Carboniferous rocks from 318 to 299 million years ago and are very similar to the most primitive surviving suborder, the Mesothelae. The main groups of modern spiders, Mygalomorphae and Araneomorphae, first appeared in the Triassic period, more than 200 million years ago.

The species Bagheera kiplingi was described as herbivorous in 2008, but all other known species are predators, mostly preying on insects and other spiders, although a few large species also take birds and lizards. An estimated 25 million tons of spiders kill 400–800 million tons of prey every year. Spiders use numerous strategies to capture prey: trapping it in sticky webs, lassoing it with sticky bolas, mimicking the prey to avoid detection, or running it down. Most detect prey mainly by sensing vibrations, but the active hunters have acute vision and hunters of the genus Portia show signs of intelligence in their choice of tactics and ability to develop new ones. Spiders' guts are too narrow to take solids, so they liquefy their food by flooding it with digestive enzymes. They also grind food with the bases of their pedipalps, as arachnids do not have the mandibles that crustaceans and insects have.

To avoid being eaten by the females, which are typically much larger, male spiders identify themselves as potential mates by a variety of complex courtship rituals. Males of most species survive a few matings, limited mainly by their short life spans. Females weave silk egg cases, each of which may contain hundreds of eggs. Females of many species care for their young, for example by carrying them around or by sharing food with them. A minority of species are social, building communal webs that may house anywhere from a few to 50,000 individuals. Social behavior ranges from precarious toleration, as in the widow spiders, to cooperative hunting and food-sharing. Although most spiders live for at most two years, tarantulas and other mygalomorph spiders can live up to 25 years in captivity.

While the venom of a few species is dangerous to humans, scientists are now researching the use of spider venom in medicine and as non-polluting pesticides. Spider silk provides a combination of lightness, strength and elasticity superior to synthetic materials, and spider silk genes have been inserted into mammals and plants to see if these can be used as silk factories. As a result of their wide range of behaviors, spiders have become common symbols in art and mythology, symbolizing various combinations of patience, cruelty and creative powers. An irrational fear of spiders is called arachnophobia. (Full article...)

Selected article on the Cenozoic in human science, culture and economics

A pick axe sitting on a wider, protruding rock. Various sized rock prudences are in the photo; most are a little wider than the pick handle. — Outcrop of Ordovician kukersite oil shale, northern Estonia

Oil shale geology is a branch of geologic sciences which studies the formation and composition of oil shales–fine-grained sedimentary rocks containing significant amounts of kerogen, and belonging to the group of sapropel fuels. Oil shale formation takes place in a number of depositional settings and has considerable compositional variation. Oil shales can be classified by their composition (carbonate minerals such as calcite or detrital minerals such as quartz and clays) or by their depositional environment (large lakes, shallow marine, and lagoon/small lake settings). Much of the organic matter in oil shales is of algal origin, but may also include remains of vascular land plants. Three major type of organic matter (macerals) in oil shale are telalginite, lamalginite, and bituminite. Some oil shale deposits also contain metals which include vanadium, zinc, copper, and uranium.

Most oil shale deposits were formed during Middle Cambrian, Early and Middle Ordovician, Late Devonian, Late Jurassic, and Paleogene times through burial by sedimentary loading on top of the algal swamp deposits, resulting in conversion of the organic matter to kerogen by diagenetic processes. The largest deposits are found in the remains of large lakes such as the deposits of the Green River Formation of Wyoming and Utah, USA. Oil-shale deposits formed in the shallow seas of continental shelves generally are much thinner than large lake basin deposits. (Full article...)

Selected images

Image 1Stegotetrabelodon is an extinct genus of primitive elephant with gomphothere-like anatomical features from the Late Miocene to Early Pliocene of Africa and Eurasia
Image 2Platybelodon ("flat-spear tusk") was a genus of large herbivorous mammals related to the elephant (order Proboscidea). It lived during the late Miocene Epoch in Asia and the Caucasus.
Image 3Victoriapithecus macinnesi was a primate from middle Miocene central Africa
Image 4Proconsul is an extinct genus of primates that existed from 23 to 25 million years ago during the Miocene epoch
Image 5Palaeotragus ("ancient goat") is a genus of very large, primitive, okapi-like giraffid from the Miocene of Africa, Eurasia.
Image 6Adcrocuta is an extinct genus of terrestrial carnivore in the family Hyaenidae that lived during the Miocene period
Image 7Indarctos is a genus of mammals of the bear family, Ursidae, endemic to North America, Europe and Asia during the Miocene
Image 8Morotopithecus bishopi is a species of fossil ape discovered in Miocene-age deposits of Moroto, Uganda.
Image 9Fossil tooth of Hyainailurus, an extinct genus of hyaenodont mammal of the family Hyainailouridae that lived during the Miocene
Image 10Deinotherium was a large prehistoric relative of modern-day elephants that appeared in the Middle Miocene and survived until the Early Pleistocene
Image 11Prolibytherium ("Before Libya's Beast") is an extinct climacoceratid artiodactyl ungulate native to Early Miocene North Africa and Pakistan
Image 12Mimoperadectes is an extinct genus of didelphimorph marsupial from the Eocene of North America